Hesperiidae

The skippers are a worldwide family of about 3500 species that appear to be "sister" to the rest of the "true butterflies". The clubs on the tips of the antennae are usually hooked. Our California skippers fall into two or three subfamilies: the spread-wing skippers (Pyrginae), the folded-wing skippers (Hesperiinae), and the Heteropterinae.

The spread-winged skippers are generally dark brown and hold both sets of their wings open when landed. They use a wide variety of hostplants, including oaks, Ceanothus, legumes, mallows, and even saltbush. California genera include PyrgusHeliopetesErynnisThorybesEpargyreus, and Pholisora.

The folded-wing skippers have a characteristic posture when they land: the forewings are held at a 45o angle to the rest of the body while the hindwing is held open and flat. This gives them a "fighter-jet" like appearance. They are largely orange and tawny, and many have whitish chevrons on the ventral hindwing, although some genera are dark brown. All members of this group feed on grasses or grassy-like plants (like sedges and rushes) as caterpillars, and as a result, they are often called the grass skippers. California genera include HesperiaOchlodesPolitesPoanesHylephilaLerodeaAmblyscirtes, and Atalopedes.

A third subfamily, the Heteropterinae, is weakly differentiated from the other skipper subfamilies, but the lone California representative is distinctive: Carterocephalus palaemon.

Lerodea eufala

A nondescript, weedy brown skipper with small white spots, a slightly grayish hindwing underside, and an odd habit of shrugging its wings several times upon alighting. Eufala ranges from our area to central Argentina and Chile. Its status in northern California is unclear. Although there are records of strays in the mountains, it basically is confined to the Central Valley and adjacent foothills where it is rarely seen before June, increasing to maximum abundance in autumn before disappearing around Thanksgiving. There is a handful of widely-scattered April records.

Ochlodes agricola

The newly-coined "common name" Rural Skipper is very unfortunate since there is a skipper "really-named" ruralis - Pyrgus ruralis. And it's in our fauna. The Farmer is intolerant of civilization. It is common to abundant in rocky foothill canyons. At mid-elevation (Washington, Lang) it occurs in dry, rocky sites, often with Goldencup Oak and Coyotemint. It does not venture above 5000'.

Ochlodes sylvanoides

Recorded from all sites; very common except at Castle Peak, where it may be only a stray. This skipper is only slightly less intolerant of civilization than O. agricola, but it occurs in a vast array of habitats, including chaparral, riparian woodland, montane coniferous forest and shrub-steppe. It flies in the second half of the season, beginning just when O. agricola ends. At low elevation it appears to have two back-to-back broods, flying from late June or early July through October. At washington and above it has only one.

Ochlodes yuma

A very distinctive, large, lightly-marked skipper found disjunctly in the southern Great Basin and southwest deserts and in the Sacramento-San Joaquin Delta and Suisun Marsh. Rare strays have turned up elsewhere; this is an extremely strong flier. The odd distribution has led to speculation that the Delta-Suisun populations may have been introduced. The oldest record seems to be from near Modesto in 1938. But there has been so little collecting in the Central Valley that that fact is not, in itself, particularly surprising.

Pholisora catullus

Perhaps we should change the common name to "The Formerly-Common Sooty Wing," since this little glossy-black butterfly with a white "face" is now teetering on the edge of regional extinction. On our transect, it was formerly found at all the low-elevation sites but as of 2005 was still present only in West Sacramento. As recently as a decade before this was a "junk species," breeding in vegetable gardens and vacant lots on weedy Pigweeds (genus Amaranthus) and occasionally on the closely-related Cockscomb in flower gardens.

Poanes melane

Although common in parts of the Bay Area where it is an urban "lawn skipper," on our transect this is entirely a species of riparian forest and is generally uncommon or even rare. It perches in dappled light and shade along streamsides, generally well off the ground. Its upper limit of residency at the latitude of I-80 seems to be about 3000'. There is no evident variation.

Polites sabuleti sabuleti

This is the multiple-brooded, somewhat seasonally-polyphenic low-altitude member of the complex that usually breeds on the turfgrass Distichlis spicata (occasionally on Bermuda Grass in lawns). It is common at Suisun, West Sacramento and North Sacramento and may colonize briefly elsewhere at low elevation. Specimens appearing to be this entity also turn up sporadically at Sierra Valley. Males perch low, often on the ground, and are highly territorial. Both sexes visit low flowers, especially Heliotrope and Lippia.

Polites sabuleti ssp.

This entity is neither the lowland P. s. sabuleti nor the high-montane P. s. tecumseh. It is marked as heavily as tecumseh (though the female ground color is generally paler here) and is as large as summer P. s. sabuleti, but with numerous pattern differences.

Polites sabuleti tecumseh

The Polites sabuleti complex may contain several genetic species. Tecumseh is the "high Sierran" subspecies, occurring from the vicinity of Bucks Lake south at mostly high altitudes and extending above tree-line in the south. It is small and heavily marked both above and below, but there is significant phenotypic and molecular variation among populations. The population at Donner Pass is typical in being univoltine and emerging in June-July (7000'). It can be quite abundant.

Polites sonora

Common at all our mountain sites, from Lang Crossing to Sierra Valley. A species of wet meadows and streamsides, usually associated with the Greenish Blue (Plebejus saepiolus) and the Nevada Cloudy-Wing (Thorybes mexicana nevada). Perches on or near the ground; visits Aster, Goldenrod, pink pussy-paws and a great variety of flowers in bloom during its flight season. There is one brood, June-August, usually peaking early. The larval host plants are undetermined grasses.