The taxonomy of this entity has undergone some very unfortunate turbulence recently. It used to be put in the species Hesperia comma, and its subspecifixc name used to be yosemite. The name harpalus was used for the East slope subspecies, which is very different. (Got that?) As now construed, harpalus refers to the single-brooded entity with a very lightly-marked, mostly golden ventral hindwing in the male and a highly variable spot-pattern on the same wing in the female, incorporating a c-shaped element on the basal half.

Until recently, this was known as Hesperia comma harpalus. Now the name harpalus has been switched to the very different West slope entity (formerly called yosemite) and most North American populations have been removed from species comma to colorado. At any rate, this insect is common along the Sierran East slope, including Sierra Valley, and on into Nevada. It looks very unlike the West slope animal.

Populations of Hesperia colorado (as determined by the genitalia) occur at scattered locations on the Sierran west slope on serpentine substrates and fly in autumn - mid-September into October. One of these occurs on serpentine along Washington Road, where it is sandwiched between "normal" populations of what is now called H. colorado harpalus but used to be called H.comma yosemite, which fly in June-July both higher and lower but on metasedimentary substrates!

The Columbian Skipper is found mainly in the Coast Range. In the Sierra Nevada it is known from only a handful of localities, mainly on serpentine. None is on the transect. Its only population on the transect is in the chaparral at the head of Gates Canyon; individuals are encountered only sporadically on the sampling route, but if the route included any rocky summits it would be a "regular" since it is a confirmed hilltopper. It also occurs in Lagoon Valley Regional Park, where it hilltops along the crest of the Cement Hill range. There are two broods, in May-July and September-October.

A striking, large skipper, like a very big East-slope Colorado Skipper with a strongly greenish tinge to the body pelage and hindwing beneath; a very strong flier; males perch on or near the ground. Although strays are occasionally taken at low altitudes, this is quintessentially a montane species, from 2000' to tree-line. It flies shortly after snowmelt (early June at Donner, e.g., visiting early Dandelions and Pussy Paws) and again, much more commonly, in autumn (August-October, mostly at Aster and Rabbitbrush).

Our populations have been named as a weakly-distinguished subspecies, H. lindseyi eldorado. An uncommon butterfly largely restricted to serpentine grassland, usually in places where there is a big display of Clarkia blossoms in late spring. This species is often sympatric with H. colorado, which it strongly resembles, but both colorado entities (q.v.) in our areea fly later in the year--one of them, much later.

In our area a high-altitude species, generally found in cold, windwept sites where males hilltop. Both sexes visit puddles and can often be seen on the wet soil adjacent to melting snowbanks. At Donner confined to the east end of the pass; on Castle and Basin Peaks and on rocky knobs along the ridge in between. This species appears to have decreased in abundance in recent years and is no longer seen every year on the transect. It usually flies in July but there are a few records as late as September. The hosts are presumed to be perennial bunchgrasses.